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There are a few key characteristics which
distinguish ferns from other plants. The most obvious is the
absence of flowers, fruits and seeds. Ferns reproduce by spores.
Spores are typically produced on the underside of the fern leaf
in clusters called sori. The original classification of the ferns
were based on the position and shape of these spore-producing
organs. The life-cycle of ferns alternates between two very distinct
generations. The fern plant, as most people are familiar with,
is the sporophyte generation and it produces dust-like spores.
Spores will germinate into minute gametophytes measuring less
than 1 centimeter. It is the gametophyte stage which produces
male sperm and female eggs and once united, the sporophyte develops. |

Polypodium fern with sori
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An emerging frond or fiddlehead
Photo by: Andrew G. Seymour
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Although mature ferns consist of roots, stems
and leaves, just as other plants do, different terms are often
used to describe the various fern parts. Fern leaves are called
fronds and the stem is called a stipe.One fairly distinctive
feature among ferns is the manner in which the newly emerging
foliage unfolds in a scroll-like fashion. The newly developed
fern leaves are called fiddleheads. Rhizomes are another distinguishing
characteristic common among many ferns. These stuctures not only
provide a vital link between the roots and the frond, but also
determines the plant's habit. Rhizomes and fronds are often clothed
with a protective covering of hairs and/or scales. Rhizomes may
be erect, holding the frond in a close, vase-like cluster, or
it may creep horizontally, with fronds arising in an irregularly. |

Salvinia auriculata - Water Spangles |

Azolla sp. - Mosquito Fern |
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There are
a number of ferns that live on or in water. These aquatic ferns
display hundreds of fine translucent hairs that hang down in
the water below them, absorbing nutrients for the plant just
as true roots. Floating ferns in their natural setting provide
cover for fish and other animals, and some are food to certain
types of fish. |
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The fern allies are relatives of the ferns.
Like the ferns, they have the alternation of generations and
they reproduce from spores. The fern allies differ mainly from
ferns in that they do not have the same leaf structure. Whisk
ferns are essentially leafless, the leaves of horsetails are
reduced to scales and the club mosses display minute single-veined
leaves.
The horsetails, called such because of
their resemblance to horses' tails, are also called scouring
rushes. The plants, which have silica deposits along the stems,
were often used for scrubbing pots and pans. The jointed stems
can be branched or unbranched, with some species reaching 15
feet. The leaves are non-photosynthetic and are nothing more
than small scales surrounding the nodes.
Club mosses and Spike mosses are, perhaps,
the fern allies which most resemble ferns. The club mosses and
spike mosses have true stems and roots, and the tiny leaves,
called microphylls, have a single vein. Selaginella, a
spike moss, branches freely, often with the branches rooting
as they creep along the soil surface. Selaginella albomarginata
is pictured at the top left of this page. |

Equisetum sp.

Psilotum nudum
Photos by: Andrew G.
Seymour |
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The Whisk Ferns, leafless shrubs about 12
inches tall, are called such because of their resemblance to
a small whisk broom. The conspicuous green plant, the sporophyte,
consists of branching stems which arise from underground rhizomes.
Photsynthesis occurs in the aboveground branching stems. No roots
are present, the rhizome performs the functions of a root. In
Psilotum, the leaves are small, scalelike outgrowths without
vascular tissue. Occasionally, small, round yellow pouches of
spores, called sporangia, appear along the branches. |
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