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Cycads are an ancient group of seed plants,
sometimes called living fossils. They first appeared in Pennsylvania
300 million years ago. In the past, cycads attained a global
distribution, extending over the earth from Alaska and Siberia
to the Antarctic. Fossils are known from every continent. In
addition to the eleven living genera, there are nineteen extinct
cycad genera known only as fossils. Many cycads are facing possible
extinction in the wild today, both because they live in endangered
habitats such as tropical forests, and because they grow so slowly
and reproduce so infrequently. |

Cycas revoluta - Sago Palm
Photo by: Andrew G. Seymour |

Cycas circinalis - Emerging cones |
Commonly called Fern Palms, cycads are neither
ferns nor palms but actually belong to the division of cone-bearing
plants known as gymnosperms. The seeds of cycads are quite large,
and are often brightly colored; red, purple, and yellow seeds
are common. These colorful seeds are displayed as the cone matures
and the seed bearing leaves separate from each other; the colors
attract birds and a variety of mammals which disperse the seeds.
Some cycads will produce heat or odors to attract animals. |

Coralloid Roots
Photo by: Andrew G. Seymour
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One of the more fascinating interactions among
these plants is an association with a photosynthetic bacteria.
In cycads, the cyanobacteria are sheltered in specially modified
roots which have the appearance of coral, and are so called coralloid
roots. These roots grow up out of the soil, rather than down
into it, and are thus exposed to light which the cyanobacteria
need. In return for providing a stable habitat, the cycad acquires
nitrogen nutrients from the bacteria. |
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The term gymnosperm comes from the greek words
gymnos, meaning naked, and sperma, a seed. This
name refers to the exposed nature of the seeds, which are not
enclosed within a fruiting body, as are the seeds of flowering
plants. There are four main divisions of gymnosperms with living
representatives: the Conifers make up the largest and most significant
division by far and are represented by pine trees, firs, spruces,
and cedars among others; the Cycads; the Ginkgoes, which have
unique fan-shaped leaves and seeds enclosed in a fleshy covering;
and the Gnetophytes, whose wood contains vessels, a structural
element typically found in flowering plants. |

Cycas circinalis |
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The Welwitschia is a gnetophyte. This cone-bearing
plant is the only one of its species and is confined to the temperate
deserts of southwestern Africa. Since there is not much yearly
rainfall, it is believed that the plant survives much of the
time on fog and dew water absorbed through the surface of the
leaves. Throughout the life of these plants, up to 100 years
or more, they only produce two straplike leaves which become
easily tattered and torn. The leaves have a meristem or growing
point at the base which constantly add to the length of the leaves.
Male and female cones are produced on separate plants, on stalks
that emerge from the axils of the leaves so that they appear
to be growing around the rim of the stem. Male cones bearing pollen are pictured in the upper
left corner of this page. |

Welwitschia mirabilis
Photo by: Andrew G. Seymour |
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